Dimetrodon

Dimetrodon (/daɪˈmiːtrəˌdɒn/ ^ⓘ or /daɪˈmɛtrəˌdɒn/; lit. 'two measures of teeth') is a genus of non-mammalian synapsid that lived during the Cisuralian age of the Early Permian period, around 295–272 million years ago. It is a member of the family Sphenacodontidae. With most species measuring 1.7–4.6 m (5.6–15.1 ft) long and weighing 28–250 kg (62–551 lb), the most prominent feature of Dimetrodon is the large neural spine sail on its back formed by elongated spines extending from the vertebrae. It was an obligate quadruped (it could only walk on four legs) and had a tall, curved skull with large teeth of different sizes set along the jaws. Most fossils have been found in the Southwestern United States, the majority of these coming from a geological deposit called the Red Beds of Texas and Oklahoma. More recently, its fossils have also been found in Germany and over a dozen species have been named since the genus was first erected in 1878.

Dimetrodon is often mistaken for a dinosaur or as a contemporary of dinosaurs in popular culture, but it became extinct some 40 million years before the advent of dinosaurs. Although reptile-like in appearance and physiology, Dimetrodon is much more closely related to mammals than to reptiles, though it is not a direct ancestor of mammals. Dimetrodon is assigned to the "non-mammalian synapsids", a group traditionally – but incorrectly – called "mammal-like reptiles", but now known as stem mammals. This groups Dimetrodon together with mammals in the clade Synapsida, while reptiles are placed in a separate clade, Sauropsida. Single openings in the skull behind each eye, known as temporal fenestrae, and other skull features distinguish Dimetrodon and true mammals from most of the earliest sauropsids.

Dimetrodon was probably one of the apex predators of the Cisuralian ecosystems, feeding on fish and tetrapods, including reptiles and amphibians. Smaller Dimetrodon species may have had different ecological roles. The sail of Dimetrodon may have been used to stabilize its spine or to heat and cool its body as a form of thermoregulation. Some recent studies argue that the sail would have been ineffective at removing heat from the body, due to large species being discovered with small sails and small species being discovered with large sails, essentially ruling out heat regulation as its main purpose. The sail was most likely used in courtship display, including threatening away rivals or showing off to potential mates.