Colubroides

The Colubroides are a clade in the suborder Serpentes (snakes). It contains over 85% of all the extant species of snakes. The largest family is Colubridae, but it also includes at least six other families, at least four of which were once classified as "Colubridae" before molecular phylogenetics helped in understanding their relationships. It has been found to be monophyletic.

Colubroides
Eastern racer (Coluber constrictor)
Scientific classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Suborder: Serpentes
Clade: Caenophidia
Clade: Colubroides
Zaher et al., 2009
Subclades
Synonyms

Xenophidia

Morphological synapomorphies are defined as such from Zaher et al. (2009):

Loss of the right carotid artery; intercostal arteries arising from the dorsal aorta throughout the trunk at intervals of several body segments; specialized expanded costal cartilages; presence of a muscle protractor laryngeus; separate muscle protractor quadrati; separate spinalis and semispinalis portion in the epaxial trunk; spinules or spines covering the hemipenial body.

Traditionally, the name "Colubroidea" was used for this clade. This was seen problematic, however, as many of the same studies that support this clade of snakes also advocated for the various subfamilies of Colubridae to be reevaluated as proper families in their own right. As the result of this, Zaher et al. (2009) proposed to rename the clade as "Colubroides", restricting Colubroidea to the clade Colubridae and several related families that were formerly colubrid subfamilies.

Below is a phylogeny of colubroid snakes found in many analyses:

Colubroides
Xenodermoidea

Xenodermidae

Colubriformes
Pareoidea
Pareidae

Pareinae

Xylophiinae

Endoglyptodonta
Viperoidea
Viperidae

Viperinae

Azemiopinae

Crotalinae

Homalopsoidea

Homalopsidae

Elapoidea

Prosymnidae

Buhoma

Psammophiidae

Pseudaspidinae

Psammophiinae

Lamprophiidae

Pseudoxyrhophiinae

Micrelapiinae

Psammodynastiinae

Lamprophiinae

Elapidae

Calliophiinae

Micrurinae

Najinae

Bungarinae

Elapsoidea

Hydrophiinae

Atractaspididae

Cyclocorinae

Atractaspidinae

Colubroidea
Colubridae

Grayiinae

Calamariinae

Ahaetuliinae

Colubrinae

Sibynophiidae

Natricidae

Pseudoxenodontidae

Dipsadidae

Carphophiinae

Xenodontinae

Dipsadinae

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