This is more a theoretical question.
Let's assume we have a Firewall/DG (with a connected WAN) on LAN IP 192.168.1.1
. All external DNS records for incoming services like OWA, FTP etc. point to this WAN.
Also on the LAN there is another Firewall/DG on 192.168.1.254
which has another WAN connection. This is the Default Gateway for the all the PCs and Servers.
In this situation - will inbound services (let's say RDP to a server, and FTP / HTTPS from the WAN connection on 192.168.1.1
which are NATTed to the servers work, given any return communication from these servers would be from the WAN address on the Firewall 192.168.1.254
(which for the sake of the question, is a totally different WAN IP and range).