Leland H. Hartwell

Leland Harrison (Lee) Hartwell (born October 30, 1939) is former president and director of the Fred Hutchinson Cancer Research Center in Seattle, Washington. He shared the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Tim Hunt, for their discoveries of protein molecules that control the division (duplication) of cells.

Leland H. Hartwell
Born (1939-10-30) October 30, 1939
Los Angeles, California, U.S.
Alma materCalifornia Institute of Technology, Massachusetts Institute of Technology
Known forCell cycle regulation
AwardsRosenstiel Award (1992)
Genetics Society of America Medal (1994)
Komen Brinker Award (1998)

Albert Lasker Award (1998)
Massry Prize (2000)
Nobel Prize in Physiology or Medicine (2001)
Mendel Medal (2001)
Medal of Merit (2003)
Scientific career
FieldsBiology
InstitutionsFred Hutchinson Cancer Research Center
Arizona State University
Biodesign Institute
Amrita Vishwa Vidyapeetham

Working in yeast, Hartwell identified the fundamental role of checkpoints in cell cycle control, and CDC genes such as CDC28, which controls the start of the cycle—the progression through G1.

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